A century and a half after On the Origin of Species, biologists still debate the mechanisms and implications (though not在《物种起源》出版了一个半世纪后，生物学家还在争论达尔文主义的机制与含意（而不是实际内容），而大多数人也还在为前达尔文时期的漫无头绪而焦躁徬徨。