Like thallose liverworts, the hornworts have an alternation of generations in which the gametophyte is a dorsiventral fl与叶状体植物地钱一样，角苔具有明显的世代交替，，配子体在世代交替中占优势，孢子体占劣势且利用其基足插入配子体的组织中吸取养料，以供孢子体生长。