The results partly correspond with the molecular evidence (Yuan & Kupfer, 99). The monophyletic Halenia clade in Yuan & Kupfer's molecular cladogram includes Halenia, S. tetraptera and Frasera (section Frasera in Swertia s.l.) ;
The results partly correspond with the molecular evidence (Yuan & Kupfer, 99). The monophyletic Halenia clade in Yuan & Kupfer's molecular cladogram includes Halenia, S. tetraptera and Frasera (section Frasera in Swertia s.l.) ;
这与Yun＆Kupfer（99）的分子证据部分吻合，因为分子系统树上的花锚单系支包括花锚、四数獐牙菜和北美的密花属（广义獐牙菜属中的密花组），而后者的胚胎学特征未有任何报道。